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ENDL. plantation. Relative radiation within and above a crown decreased exponentially with increasing cumulative leaf area density per unit crown projection area from the top of the crown downward. Middle-sized trees showed steep light attenuation as compared with large- or small-sized trees. Gross photosynthetic production on a whole tree basis was evaluated using a modified model of the Monsi-Saeki model. As the total leaf area of a tree approached a critical leaf area, the gross photosynthetic production dropped abruptly. The dependence of the gross photosynthetic production on the total leaf area was described by a generalized power function. There was no considerable difference in gross photosynthetic production per unit leaf area, or in mean photosynthetic activity at a whole tree level, among larger-sized trees accounting for 85% of the total number of trees. The gross photosynthetic production per unit leaf area decreased more drastically in the smallest-sized tree having foliage near the critical leaf area. 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